Variation in Wigeons

Every now and then I hear discussions about Eurasian Wigeon that have turned up in North America showing some green behind the eye.  Most books don't illustrate this and the sceptics cry, "Hybrid!".  Of course most of us who are lucky enough to look at lots of Wigeon this side of the Atlantic know well that Wigeon often show a little green behind the eye - it's not a sign of hybrid origin.

But what about Wigeon with complete green headbands?  Is this just normal variation?  Or are these birds hybrids?  Like these:

Eurasian Wigeon, Pentney Lakes, 31st March 2009

Eurasian Wigeon, Burnham Norton, 13th March 2010

Most people I've discussed this with consider that it's just the extreme of variation within pure Wigeons.  These birds don't show any other sign of hybridisation so they can't be first-generation hybrids, and many first-generation Eurasian x American Wigeon hybrids don't show a green headband like this anyway.  The theory is that Wigeons have inherited the mix of genes required to produce a green headband from a common ancestor but that these genes are not usually expressed in Eurasian Wigeon.  Always present, normally suppressed, but expressed in just a few individuals.  A recent discussion on ID-Frontiers seemed to reach the same conclusion but thanks to Joern, Carl Gunnar, Lars and Henry with whom I had already discussed these birds.

Whilst this theory has much merit, two observations haven't sat comfortably with it for me:

1. My experience was that Wigeon with a little green behind the eye are common and Wigeon with an extensive green band are occasional, but I'd never noticed any Wigeon with anything in between.  If it were simply variation I'd expect a continuum, with the greater extents being increasingly rare. 
2. I've heard one or two people with experience of Wigeons in Europe and in the Far East claiming that birds in the Far East have a much higher tendency to show green head bands.  American Wigeons turn up in the Far East more often than in Europe and so backcrossed hybrids might be expected more commonly there than here.  Moreover someone recently claimed that Wigeon in India (where so far as I know American Wigeon has never been recorded) never show any green behind the eye.  Thus anecdotal evidence seems to suggest that the occurrence of green on the head correlates with the frequency of vagrant American Wigeons in the population.

Well this is one mystery I made some progress with recently.  I had the opportunity to study a few hundred Wigeon at close range and in excellent light the other day.  The main thing I learned is that there is in fact much more variation than I had appreciated, and it's more common than I'd realised.  My initial observations described as point 1 above proved not to be valid - there is indeed a continuum, with many birds showing green extending further behind the eye than most, but not as far as the few.

Eurasian Wigeon, Snettisham, 8th December 2012

It still doesn't explain the geographic variation but there may be other explanations for that.  Moreover it's only anecdotal so far as I know and like my first point, maybe it won't stand up to more careful observation.  Sadly I've got no trips to the Far East or India planned so I won't be able to test that one myself - but if you travel more widely let me know if it stacks up with your experience!

I noticed a couple of other things on some of the earlier green-banded Wigeons I saw, which I wondered if they might be linked.  A couple showed a small dark (green?) wedge at the upper point of the cream forecrown, absent from most Wigeon.  However I've since found this on other Wigeon that don't have green behind the eye, so it seems not to be linked.  One also showed pale at the base of the bill extending further into the cheek area than normal, but again this seems to be a variable feature and not linked to the extent of green behind the eye.

Eurasian Wigeon, Pentney, 31st March 2009 

(the same bird as the one at the top of this post, now showing the dark wedge at the apex of the cream forecrown)

Nothing to do with the above, but worth a mention while I'm talking about variation in Wigeon, I also had the opportunity to photograph a few first-winter drake Wigeon in flight.  It's well-known that young male Wigeon lack the white forewing of adults but perhaps not quite so well-known that they continue to lack this after they've moulted their head and body feathers.  Actually it's quite well documented, so this is nothing new, but I thought it was interesting to show a couple of shots that illustrate it nicely.

first-winter male Eurasian Wigeon, Snettisham, 8th December 2012

 

Lots more photos of Wigeon on my website of course.

And while we're talking about green on duck heads where we perhaps don't expect it, here's a nice first-winter drake Pintail with a green sheen.  Not the same issue as the Wigeon here of course - in this instance I think it's just a case of how the light catches it, a bit like Tufted Duck I suppose only less obvious.

first-winter male Pintail, Snettisham, 8th December 2012

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Follow up, Feb 2013

A quick postscript following more study of Wigeon over the course of the winter.  I knew that the green on a duck's head is irridescent, i.e. the intensity of colour and the colour itself is variable according to the intensity of the sunlight and the bird's angle to the light compared to the observer.  I had even noticed that some of my observations of Wigeon with the most green on the head had been in strong sunshine (including the photos above).  But I hadn't quite realised the extent to which the light impacts the colour when the normal ground colour is chestnut-brown as on a Wigeon's head.

A few days after I posted this I studied a flock of several hundred Wigeon a bit further round the Norfolk coast in dull and cloudy conditions - not one of them showed the slightest hint of green in their head.  

Many of the questions about variability remain.  Most Wigeons don't seem to show green behind the eye even when it's sunny.  A few do even when it's not sunny.  What causes that variation is still an unanswered question, but the impact of sunlight does mean it will be difficult to assess how much it varies geographically.  As the extent of green varies according to the brightness and direction of sunlight it will be nigh-on impossible to make meaningful comparative assessments between different populations.

Thanks to everyone who has commented privately about this subject, and especially for the offers to get involved in further study in different countries, were we to find a meaningful way of doing that.

Another non-juvenile immature skua

Following my last post about a first-summer Pomarine Skua here's another immature skua that's not a juvenile.  This one was seen on a sea watch from Sheringham in Norfolk.

I'm not completely sure if it's first-summer or second-summer.  Whilst it had a dark forehead and forecrown covering pretty much the same area as the dark on an adult pale-phase Arctic Skua it wasn't at all well-defined and didn't really recall the dark cap of an adult.  From what I've read a second-summer would be better defined than this, but do you agree that it must therefore be first-summer?  Are there any other pointers visible in the photos that enable us to age it?

And what about the species.  Having discussed it with Rob I think it's an Arctic, and that's what I thought it was in the field too.  While I was watching it I did wonder if Long-tailed could be ruled out, and I'm still not completely certain I've adequately eliminated that possibility.  What do you think?  Is there anything in the photos that clinches it beyond doubt?

probable first-summer Arctic Skua, Sheringham, 27th September 2012

First-summer Pom

I was hoping to come back from the Scilly birders' pelagic with photos of Wilson's Petrels to share with you but sadly that didn't quite work out (again).

Instead here's something a bit different - a Pomarine Skua in first-summer plumage.  Apparently they don't normally get many Poms down there but this year there have been a few, nearly all first-summers.  Not a plumage we get to see very often here in Norfolk, although if I'm perfectly honest it's quite possible I've overlooked it in the past.

Pomarine Skua, MV Sapphire birders' pelagic trip, 12th August 2012

Oh, and while I'm at it I might as well show off a bit of Great Shear action... something else we don't see in Norfolk very often.

Great Shearwater, MV Sapphire birders' pelagic trip, 12th August 2012

Confusing Yellow Wagtails - part 2

One enjoyable moment hunting for migrants in Sicily was finding a mixed flock of 10 flava Wagtails in a dried out pool at Capo Murro di Porco.  It contained Blue-headed Wagtails, Grey-headed Wagtails, a Black-headed Wagtail (nice call) and another curious bird, the point of this post.

Before I get on to the bird in question, let me wallow in some Grey-headed loveliness:

Grey-headed Wagtail, Capo Murro di Porco, 4th May 2012

I didn't manage to get such a good shot of the Black-headed, but it was such a gorgeous bird I can't resist sharing it:

Black-headed Wagtail, Capo Murro di Porco, 4th May 2012

And now, the bird I'm really writing about.  This was a really striking bird!  A lovely dark slaty-grey head, really dark at the front (fore-crown, lores and front of ear-coverts) but paler, almost bluey grey at the rear.  A vivid white supercilium and white lower eye-crescent, but no white in the middle of the ear-coverts.  The throat was clearly yellow but the chin was white, with white extending slightly down the moustaches.  Here it is:

flava Wagtail, Capo Murro di Porco, 4th May

So what is it?  Yellow throat should rule out the cinereocapilla/iberiae complex  I thought it might be a Romanian 'dombrowski'-type intergrade?  What do you think?

Confusing Yellow Wagtails - part 1

I was hoping to score some nice Ashy-headed Wagtails in Sicily, and so I did.  Only at one site I visited though - the wetlands at Granelli.  Only managed to get one distant shot of a 'textbook' Ashy-headed though:

Ashy-headed Wagtail, Granelli, 1st May 2012

One bird was much closer, and as it was singing its heart out and clearly holding territory it should have been Ashy-headed... but what about that supercilium?

flava Wagtail holding territory, Granelli, 3rd May 2012

Now, is it me, or does this look rather like a Spanish Wagtail iberiae?  Are there Spanish Wagtails breeding in Sicily?  I saw iberiae in Portugal a few years ago and I think most of them had a narrower supercilium in front of the eye, but variable.  Or is it an intergrade/hybrid (with Blue-headed?)?  Or can Ashy-headed Wagtail look like this?  What would you think it was if you saw it in England?


Here's another.  This one was in the same area and looked more like Blue-headed, showing both a pale supercilium and pale in the middle of the ear-coverts.  But the tone of grey on the head - that didn't look like Blue-headed.

flava Wagtail, Granelli, 28th April 2012

Both of these birds have some green on the sides of the crown.  What does that indicate?  I've seen it on Blue-headed Wagtails previously but never quite figured out if it's an age-related thing or something else.  I wouldn't have any trouble seeing the second bird above as a first-summer - when I first glimpsed it I thought it was a female.  But the strikingly colourful first bird - can that be immature?


So help me out - what are we dealing with here?